The teams on the two worlds work together to solve the case of a shapeshifter, and Agent Lincoln Lee joins his world's team when no one remembers the existence of Peter Bishop.
A mysterious millionaire, Finch, approaches John Reese, a CIA field officer who is believed dead, and offers him the chance to stop violent crimes before they happen.
After capturing an Alpha working as an obstetrician, the team takes on an Alpha with the ability to remain unseen and undetectable against any opponent.